Each of these things has three real-valued "genes", phi, beta, and chi. Phi is sort of a generic "fitness" attribute. It's used to select which of the things of gender I is going to reproduce in the current step. Chi influences the mate selection process described above. If it's positive, there is a preference for high Beta in the mate. If it's negative, there is a preference for low Beta. If it's 0, neutral. Beta, therefore, obviously influences the rate of a gender-II critter's chance of being selected as a mate, and also it as the effect of decreasing the effective value of Phi above. Beta is the "showing off" or "costly signalling" parameter.
Ideally, if this model were to effectively confirm the theory